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AAEAAAAAAAABAgAAAAEAAAAAAAABAwAAAAEAAAAAAAABBAAAAAEAAAAAAAABBQAAAGhuYW1kAAAA YAAAAAIAAAAEAAAAAAAAAAMAAAAAAAAACgBBAHIAaQBhAGwAAAAAAAgAAAAAAAAAAwAAAAAAAAAm AE0AbwBuAG8AdAB5AHAAZQAgAFQAeQBwAG8AZwByAGEAcABoAHkAAAAAAQYAAAAEAAwAAAAAAQcA AAAGAAAAAAAAAAAABAAAAA4ACQARAAAAGgABAAAACAwBAAAAAAACAAABDAAAAAAAAAAUAAAAIAAA AAEAAAABAAAAAAAAAAEAAADoAAAACAAAAAQAAAAAAAABAAAAAAEAAAAAAAABAQAAAAEAAAAAAAAB AgAAAAEAAAAAAAABAwAAAAEAAAAAAAABBAAAAAEAAAAAAAABBQAAAGhuYW1kAAAAYAAAAAIAAAAE AAAAAAAAAAMAAAAAAAAACgBBAHIAaQBhAGwAAAAAAAgAAAAAAAAAAwAAAAAAAAAmAE0AbwBuAG8A dAB5AHAAZQAgAFQAeQBwAG8AZwByAGEAcABoAHkAAAAAAQYAAAAEAAwAAAAAAQcAAAAGAAAAAAAA AAAABAAAAA4ACQARAAAAGgABAAAAAAAAC/AEAAAAEjRWeD== ------=_NextPart_01C8AF8E.59472230 Content-Location: file:///C:/E1C9598E/Myosin_files/slide0002.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" The length of the myosin neck determines the step size
The actomyosin interaction
=
=
=
=
ADP
ADP.Pi<= /span>
ATP
Pi<= /span>
ADP
ATP
1
4
3
2
1. The rigor complex. No nucleotide bound to myosin. Tight binding between actin and myosin
2. ATP binds to myosin, head dissociates from actin
3. ATP hydrolyzed to ADP.Pi, lever arm cocks
4. Myosin:ADP.Pi binds to actin, Pi is released, lever arm swings, actin and myosin filaments slide relative to each other. ADP dissociates
How is the hydrolysis of= ATP translated into movement of the lever arm?
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iYcCABYAA4mDAQQYHh8gIQWRAAEAIigpKisZBpEHGiwtODk6LhCRBCMvO0RFRjwbkQkjMD1HSEk+ MQmKACw/SktMQDIKnYQBEyQzQU1OQjQKxSU0P0FAPSYLz9AUIjVDPDYSpZEMJx43KBgNm4YRHRUJ 7faCDv2DAgEAOw== ------=_NextPart_01C8AF8E.59472230 Content-Location: file:///C:/E1C9598E/Myosin_files/slide0004.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" The length of the myosin neck determines the step size
Myosin motor domain structure
Nucleotide
Relay = helix
Lower 50 K domain (actin binding domain)
Convert= er domain
(green)=
Note t= he cleft between the upper 50K domain (red) and the actin binding domain (grey).
The ac= tin binding cleft exists in an open conformation (weak actin binding) and a closed conformation (strong actin binding)
The nu= cleotide binding domain is
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Content-Location: file:///C:/E1C9598E/Myosin_files/slide0007.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" The length of the myosin neck determines the step size
The actin binding cleft exis= ts in both open and closed conformations:
=
Open: nucleotide bound, weak actin binding
Closed: no nucleotide bound, strong actin binding
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